Since it occurred in only one county (Douglas, the northwesternmost),

we also provide results for lowland roadsides excluding this species (Table 3). In that case, the remaining specialist species were similarly abundant in bogs and lowland roadsides, but consistently decreased in proportion of total butterfly individuals from bog to lowland roadside to upland roadside. Total butterfly abundance was much lower in bogs, and similarly higher in lowland and upland roadsides (Table 3). Table 3 Mean, minimum, and maximum relative abundance (observation rate of individuals/h) BI 10773 cell line of each species group and total individuals (including unidentified individuals), and proportion (%) of each species group out of total individuals, per year during 2002–2009   Specialists Affiliates Generalists Immigrants Total Rate % Rate % Rate % Rate % Rate Bog  Mean 21.6 44.90 18.2 34.00 9.3 18.20 1.4 2.90 54.3  Minimum 15.9 24.90 5.2 15.30 4.9 10.70 0.1 PF299804 research buy 0.20 32.4  Maximum 29.9 68.00 30.8 52.80 22.8 35.80 5.5 9.30 74.5 Lowland roadsides  Mean 51.3 33.00 24.9 17.90 68.2 48.30 1 0.70 149.3  Minimum 20.2 15.50 14.3 7.70 35.6 34.40 0.2

0.10 106.4  Maximum 140.7 56.40 47.8 27.60 97.6 63.10 2.9 2.30 255.9 Lowland roadsides (excluding Boloria montinus)  Mean 22.7 18.50             120.7  Minimum 3.8 4.30             78.6  Maximum 63.1 36.70             178.1 Upland roadsides  Mean 0.2 0.20 10 8.00 121.8 88.10 3.8 3.70 138.4  Minimum 0 0.00 4.7 2.70 42.4 78.00 0 0.00 49  Maximum 0.7 0.90 21.3 13.40 257 95.10 14.1 13.80

286.2 We recorded the same bog specialist and affiliate species in muskegs as reported in Nekola’s (1998) study; additional species we recorded in kettleholes and coastal peatlands within Nekola’s (1998) study region were infrequently encountered in only one or two sites per bog type (Table 4). Table 4 Presence of the ten peatland species analyzed by Selleckchem Ruxolitinib Nekola (1998) in the three bog types   Muskeg Kettlehole Coastala Bog specialists  L Lycaena epixanthe N S/S N S/S N S  L Lycaena Depsipeptide in vitro dorcas N S/S       S  N Boloria freija N S/S   S/   S  N Boloria frigga N S/S          N Boloria eunomia N S/S N S/S N S  N Boloria montinus N S/          N Erebia discoidalis N S/S          N Oeneis jutta N S/S N S/S   S Bog affiliates  L Callophrys augustinus N S/S N S/S N S  N Coenonympha tullia N S/S N S/S N S By study reporting them: N reported by Nekola and S reported by this study in the northwest/northeast subregions; all Swengel additions are in kettleholes and coastal peatlands within Nekola’s study region Species occurrences in this study in bog types where they were not reported by Nekola (1998): Kettlehole 1 B. freija in East Wishbone Lake each in 3 years (all attempts in that unit).