lactinea) Regularly pored becoming daedalean to lamellate never Artolenzites Glabrous-dull None Sordid yellow Contracted into a stem-like base – sometimes with a disc Pored, daedalean to lamellate often in a single specimen-irregular never T. ljubarskyi-T. cingulata Glabrous-dull to semi glossy Colorless, becoming black with KOH 5% for T. cingulata Deep brown (T. ljubarskyi)

to strongly black (T. cingulata) Never contracted into a stem-like base Regularly pored never L. warnieri Glabrous-dull none Context pale brown-abhymenial surface deep brown Never contracted RG-7388 chemical structure into a stem-like base Regularly lamellate never This classification is nevertheless incomplete, since some critical taxa from various tropical parts of the world were not accessible to us and might either add
ages to the system, or illustrate more continuities between some of the proposed divisions. In the same way two still unplaced lineages not included in previous analyses: ‘Lenzites’ warneri and the ‘Trametes’ ljubarskyi-T.

cingulata group, cannot reasonably justify new genera according to their uncertain position in our analyses, Adavosertib nor can they be included in Trametes s.s. because of outstanding morphological features, and will deserve further studies. There are here provisionally maintained in their traditional genera. Morphological characters in the four branches within the Trametes clade Structure of upper surface Aspect and structure of the abhymenial surface is a discriminating morphological feature

of major importance at the generic level in the core polyporoid clade, as already shown in Ganoderma (Steyaert 1980; Gottlieb et al. 1999; Moncalvo 2000; Welti and Courtecuisse 2010). In the Trametes group differences in pileus-structure (glabrous or tomentose) have already been described for each species studied here new and are considered by Læssøe and Ryvarden (2010) as an essential feature for species recognition; they nevertheless never been used for phylogenetic interpretation. Taking our phylogenetic results, fundamental differences in structure (Fig. 4) and consequently in macroscopic aspect of the basidiome surface, explain the evolutionary history of the groups. Differentiation of hairs (pileus tomentum) is a synapomorphy of our redefined genus Trametes (Fig. 4a–c), without any known exception, although some species are only minutely pubescent when young and become somewhat glabrous buy CP673451 whilst ageing (T. gibbosa, T. ochracea, T. suaveolens). Fig. 4 Pileus structures in Trametes and allied species. a: trichoderm with differentiated subpellis, with incrustations (Trametes versicolor); b: idem, without incrustations (T. villosa); c: trichoderm without differentiated subpellis (T.